Dire Wolf


Early in 2025 American news media pounced on a story: The Return of the Dire Wolf. Details of this remarkable piece of journalism are explained in a remarkable Wikipedia article: Dire wolf. Be sure to see the final section of the article.

illustration of dire wolf
Dire Wolf, Courtesy of paleoartist Mark Hallett.

Actually, real dire wolves roamed America from about 250,000 years ago until about 10,000 years ago. The image shown here, proportioned to precisely measured skeletal remains, is probably as accurate as any ever drawn. The dire wolf looked much like the modern grey wolf but was about 25% larger. I thank Mark Hallett for permission to show this image. Hallett is well known for his work in paleoart. Among his many accomplishments, he was chosen by Stephen Spielberg to help create the dinosaurs in Jurassic Park. See Mark Hallett: Capturing the Essence of Prehistoric Life.

The Evansville Dire Wolf

The type specimen for the dire wolf (Aenocyon dirus - a mix of Greek (dreadful + dog) and Latin (fearsome) was found in Evansville, Indiana. Late in the summer of 1854 Francis A. Linck found the specimen in the bed of the Ohio River. At that time, the depth of the river averaged about eleven feet lower than today. Linck found the specimen near the mouth of Pigeon Creek and a short walk from downtown Evansville. Although Linck refused to let Dr. Norwood of nearby New Harmony send the fossil to Dr. Joseph Leidy in Philadelphia, he died on 28 August 1854, and his family granted Dr. Norwood's wish. Dr. Leidy determined that the fossil represented a new species of wolf, and the name - Canis dirus Leidy - became official in 1858, until, in 2021, it was changed to Aenocyon dirus.)

type specimen of dire wolf
Evansville waterfront, 1856, including bridge over Pigeon Creek and banks of
the Ohio River, where the type specimen of the dire wolf was found.
Courtesy of Willard Library, Evansville.
type specimen of dire wolf
The type specimen of Aenocyon dirus (dire wolf), with notes indicating an earlier name referring to Indiana.
Courtesy of Academy of Natural Sciences of Drexel University.

It was my good fortune to find that Dr. Norwood's letters to Dr. Leidy have survived in Philadelphia. I obtained copies and found Dr. Norwood's handwriting neat, heavy, and easy to read. Dr. Norwood, it turned out, was the first state geologist of Illinois, and he worked closely with Dr. Owen, also of New Harmony, who was the first state geologist of Indiana. For an article about these two medical doctors who became geologists, beginning with an account of the Evansville dire wolf; see "David Dale Owen and Joseph Granville Norwood: Pioneer Geologists in Indiana and Illinois".

Joseph Norwood
Joseph Norwood. Photograph reproduced from George P. Merrill, The First One Hundred Years of American Geology (New Haven, Conn., 1924). Signature from Norwood's letter of acceptance as Correspondent of the Academy of Natural Sciences of Philadelphia, May 23, 1843.
Joseph Leidy
Joseph Leidy, 1859.
Courtesy of Academy of Natural Sciences of Drexel University.
Norwood letter to Leidy
Dr. Norwood to Dr. Leidy, 12 August 1854 pages 1 and 4.
Courtesy of Academy of Natural Sciences of Drexel University
Norwood letter to Leidy
Dr. Norwood to Dr. Leidy, 12 August 1854 pages 2 and 3.
Courtesy of Academy of Natural Sciences of Drexel University

The drawings of bones found by Linck, made by John Chappellsmith are not mentioned in Leidy's publications. It appears that they no longer exist. Although Henry Pratten, is mentioned in Norwood's letter, he appears to have had not recorded connections with Linck's findings. Pratten was Norwood's assitant in New Harmony and Illinois. He is mentioned in Norwood's letters to Leidy on a variety of subjects, both before and after the letter that appears here.

Earliest publications

The first publication to describe the "new" extinct species was written by Joseph Leidy, M.D. It appeared in the Proceedings of the Academy of Natural Sciences, 1854-1855, vol. 7, pp.195-201, dated November 1854, under the heading Notice of some Fossil Bones discovered by Mr. Francis A. Lincke [sic], in the banks of the Ohio River, Indiana. "The bones are usually found in this locality sticking in the bank, when the water in the river is low." Leidy continues, "The collection contains bones of the following animals:—:" Megalonyx Jeffersonii [giant ground sloth, extinct], Bison Americanus (?), Cervus Virginianus (common American deer), Equus Americanus (American horse, extinct), Tapirus Haysii (tapir, extinct), and Canis Primævus. It is with the latter name that Leidy introduces the species now known by the common name dire wolf. He recognizes that other naturalists could assert that Canis Primævus is within the compass of the previously established species canis lupus. He also devotes a paragraph to the general question of origin of species, "whether from a single pair or from a plural stock, whether by ascent from the lowest for of life, in gradually diverging series ...", an interesting digression published five years before Darwin's On the Origin of Species. Leidy concludes the article with a table of eight dental measurements from five species of Canis; in all eight, the measurements for Canis Primævus. exceed the others.

The second publication, which cites the first, appears in Journal of the Academy of Natural Sciences. Here, Leidy repeats the content of the Journal article and adds a few details and notes regarding the origin of species. His third publication, in an article, "Notice of remains of extinct vertebrata, from the Valley of the Niobrara River...[1857], Proceedings of the Academy of Natural Sciences, comments, at the end of a discussion of a different species of wolf found in Nebraska, that "the name canis primævus was inadvertently applied" [because another species had already been given that name], and that the name should be changed to canis dirus. This taxon survived for more than 160 years, to be replaced in 2021 by the present taxon, Aenocyon dirus.

Dire wolves at the La Brea Tar Pits

This section pending. Will include this reference: "The Asphalt Group of Fossil Skeletons: The Tar-Pits of Rancho-La-Brea, California, by W. D;. Matthew, in The American Museum Journal, vol. XIII, No. 7, Nov 1913, pp. 291-297.
Scene at La Brea tar pits
Scene at La Brea Tar Pits: two dire wolves, saber-toothed cat, carcass of Columbian mammoth.
Painted by Robert Bruce Horsfall, 1911
Scene at La Brea tar pits
Dire Wolf, by Erwin Sachem Christman, 1913
Mural of La Brea Tar Pits
Mural of La Brea Tar Pits, by Charles R. Knight, 1921
Mural of La Brea Tar Pits
Mural of La Brea Tar Pits, by Mark Hallett, copyright 1988.
Used by permission.
Mural of La Brea Tar Pits
Dire wolves in Mark Hallett's 1988 mural.
Used by permission.

Dire wolves, worldwide

In American popular culture, the dire wolf is a menacing creature with piercing eyes and steely fangs, lurking in the darkness and in the song, "Dire Wolf", by Grateful Dead. The more recent contribution to mythology is represented by three white "dire wolves", or genetically modified grey wolves, named Romulus, Remus, and Kaleesi.


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